Heyer WR. Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, Northern South America, and Amazonia . Arq. Zool. 2005;37(3):269-348. doi:10.11606/issn.2176-7793.v37i3p269-348