Gnaspini, P. 1994. Phylogenetic analysis of the Tribe Ptomaphagini, with description of new neotropical genera and species (Coleoptera, Leiodidae, Cholevinae, Ptomaphagini). Papéis Avulsos de Zoologia. 39, (Feb. 1994), 509–556. DOI:https://doi.org/10.11606/0031-1049.1994.39.p509-556.