R. D. MacCulloch and A. Lathrop, “Hylid frogs from Mount Ayanganna, Guyana: new species, redescriptions, and distributional records”, Phyllomedusa, vol. 4, no. 1, pp. 17–37, Jun. 2005, doi: 10.11606/issn.2316-9079.v4i1p17-37.