Description of the late- or last-instar larva of Ischyomius Chevrolat, with comments on the family placement of the genus (Coleoptera: Tenebrionoidea: Pythidae)

Authors

DOI:

https://doi.org/10.11606/1807-0205/2020.60.special-issue.11

Keywords:

Immatures, Neotropical, Classification, Zingiberales, Ecuador, Venezuela

Abstract

Study of recently collected larvae of two species of Ischyomius Chevrolat, 1871 from Napo and Pichincha, Ecuador, tentatively identified as I. denticollis Champion, 1916 and I. singularis Chevrolat, 1871 confirm the classification of the genus Ischyomius within the family Pythidae. Salient larval characters which solidify this placement are: 5 stemmata arranged into an anterior column of 3 and posterior column of 2, parabasal ridges along anterior margins of abdominal tergites, prominent urogomphi with inner urogomphal teeth and various dorsal tubercles, and transverse row of dentiform asperities along anterior margin of sternite 9. Important features of the larva are illustrated with photographs. A key to larvae of world genera of Pythidae is provided as well as natural history information on Ischyomius.

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Author Biographies

  • Darren A. Pollock, Eastern New Mexico University (ENMU) - Portales, New Mexico, United States.

    Portales, New Mexico, United States.

  • Jim McClarin, Pontificia Universidad Católica del Ecuador (PUCE), Facultad de Ciencias Biologicas, Museo de Zoología (QCAZ), Sección de Invertebrados - Quito, Pichincha, Ecuador.

    Quito, Pichincha, Ecuador.

References

Lawrence, J.F. 1982. Coleoptera. In: Parker, S.P. (Ed.). Synopsis and classification of living organisms. New York, McGraw Hill. p. 485-553.

Lawrence, J.F. 1991. Order Coleoptera (Introduction). In: Stehr, F.W. (Ed.). Immature Insects. Dubuque, Kendall/Hunt. v.2, p. 144-298.

Lawrence, J.F. & Newton Jr., A.F. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). In: Pakaluk, J. & Ślipiński, A. (Eds.). Biology, phylogeny, and classification of Coleoptera. Papers Celebrating the 80th Birthday of Roy A. Crowson. Warszawa, Muzeum i Instytut Zoologii PAN. p. 779-1092.

Lawrence, J.F.; Ślipiński, A.; Seago, A.E.; Thayer, M.K.; Newton, A.F. & Marvaldi, A.E. 2011. Phylogeny of the Coleoptera based on morphological characters of adults and larvae. Annals of Zoology, Warszawa, 61: 1-217. http://doi.org/10.3161/000345411X576725

McKenna, D.D.; Wild, A.L.; Kanda, K.; Bellamy, C.L.; Beutel, R.G.; Caterino, M.S.; Farnum, C.W.; Hawks, D.C.; Ivie, M.A.; Jameson, M.L.; Leschen, R.A.B.; Marvaldi, A.E.; McHugh, J.V.; Newton, A.F.; Robertson, J.A.; Thayer, M.K.; Whiting, M.F.; Lawrence, J.F.; Ślipiński, A.; Maddison, D.R. & Farrell, B.D. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology, 40: 835-880.

Pollock, D.A. 1994. Systematic position of Pilipalpinae (Coleoptera: Tenebrionoidea) and composition of Pyrochroidae. The Canadian Entomologist, 126: 515-532.

Pollock, D.A. 1998. Revision of the Neotropical genus Ischyomius with a discussion on its systematic position (Insecta: Coleoptera: Tenebrionoidea: Pythidae). Annals of Carnegie Museum, 67(3): 245-264.

Pollock, D.A. 2007. A new species of Ischyomius Chevrolat (Coleoptera: Pythidae) from Costa Rica. Coleopterists Bulletin, 61(2): 277-280.

Pollock, D.A. 2009. A new species of Ischyomius from Venezuela (Coleoptera, Tenebrionoidea, Pythidae) with a revised key to world species. ZooKeys, 6: 69-74.

Pollock, D.A. 2010. Pythidae Solier, 1834. In: Leschen, R.A.B.; Beutel, R.G. & Lawrence, J.F. (Eds.). Coleoptera, Beetles Vol. 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). De Gruyter, Berlin, 708-715.

Pollock, D.A. 2018. Beetles (Coleoptera) of Peru: a survey of the families. Pythidae Solier, 1834. Revista Peruana de Biología, 25(3): 321-324. http://doi.org/10.15381/rpb.v25i3.15215

Pollock, D.A. & Lawrence, J.F. 1995. Review of Anaplopus Blackburn (Coleoptera: Pythidae), with comments on constituents and systematics of Pythidae. In: Pakaluk, J. & Ślipiński, A. (Eds.). Biology, phylogeny, and classification of Coleoptera. Papers Celebrating the 80th Birthday of Roy A. Crowson. Warszawa, Muzeum i Instytut Zoologii PAN. p. 449-472.

Watt, J.C. 1987. The family and subfamily classification and New Zealand genera of Pythidae and Scraptiidae (Coleoptera). Systematic Entomology, 12: 111-136.

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Published

2020-03-04

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Original Article

How to Cite

Pollock, D. A., & McClarin, J. (2020). Description of the late- or last-instar larva of Ischyomius Chevrolat, with comments on the family placement of the genus (Coleoptera: Tenebrionoidea: Pythidae). Papéis Avulsos De Zoologia, 60(special), e202060(s.i.).11. https://doi.org/10.11606/1807-0205/2020.60.special-issue.11