Addressing nomenclatural issues in South American marine bivalves
DOI:
https://doi.org/10.11606/1807-0205/2026.66.031Keywords:
Bivalvia, Homonymy, ICZN code, Mollusca, Nomen oblitum, Synonymies, Taxonomy discussionsAbstract
This paper critically examines several taxonomic issues specific to the South American bivalve fauna, with a particular focus on the Atlantic coast. Some of these issues are introduced or endorsed by Huber’s two compendia on bivalves (2010, 2015), which led to significant taxonomic and nomenclatural changes across the global Bivalvia. Here four nomina oblita/nomina protecta cases are analyzed in this study. One of them is resolved: Phyllodina tenuisculpta (Sowerby II, 1868) is treated as a nomen oblitum, while Phyllodina persica (Dall & Simpson, 1901) is recognized as a nomen protectum. For the remaining three cases, this paper fulfills the requirements of Article 23.9.2 in asserting that status; however, a definitive resolution depends on submissions to the Commission to address pending issues to attend Articles 23.9.3 and 23.9.6. All arguments are presented in detail. The taxa are: Temnoconcha galathaea (Lamarck, 1818) should be considered a nomen oblitum, with Temnoconcha brasiliana (Dall, 1921) recognized as valid and nomen protectum. Similarly, Chionopsis crenata (Gmelin, 1791) should be also identified as a nomen oblitum, with Chionopsis pubera (Bruguière, 1791) established as valid and nomen protectum, including a rectification of authority. Spondylus tenuis Schreibers, 1793 and S. ambiguus Chenu, 1844 should be classified as both a nomen oblitum and, the former, a possible n. dubium, while Spondylus ictericus Reeve, 1856 should be accepted as valid and nomen protectum. Four synonymies are also analyzed. The synonymy of Anomalocardia flexuosa (Linnaeus, 1767) and A. brasiliana (Gmelin, 1791) is provisionally accepted, considering that they may represent distinct entities, as is the case with Tivela dentaria (Lamarck, 1818) and T. isabelleana (d’Orbigny, 1846). The synonymy between Sphenia fragilis (H. & A. Adams, 1854) and Sphenia antillensis Dall & Simpson, 1901 is rejected, with the latter updated to S. iheringiana (Pilsbry, 1897). Additionally, the synonymy between Petricolaria serrata (Deshayes, 1853) and Petricola stellae Narchi, 1975 is also disregarded. All justifications are properly explained and discussed.
Downloads
References
Abbott, R.T. 1974. American Seashells. 2. ed. New York, Van Nostrand Reinhold Co. 663p., 240 pls.
Adams, H. & Adams, A. 1854. Description of a new genus of bivalve Mollusca. Annals and Magazine of Natural History Ser. 2, 14: 418. https://doi.org/10.1080/037454809494368.
Altena, C.O.v.R. 1968. The Holocene and Recent marine bivalve Mollusca of Surinam. Studies on the Fauna of Curaçao and other Guyanas, 10(42): 153-179.
Altena, C.O.v.R. 1969. Mollusca from boring “Alliance-28” in Surinam (Dutch Guiana). Geologie en Mijnbouw, 48(2): 177-179.
Altena, C.O.v.R. 1971. The marine Mollusca of Suriname (Dutch Guiana) Holocene and recent. Part II. Bivalvia and Scaphopoda. Zoologische Verhandelingen, 119: 3-100.
Arruda, E.P.; Denadai, M.R.; Quast, M.P. & Amaral, A.C.Z. 2011. Bivalvia (pp. 236-250). In: Amaral, A.C.Z. & Nallin, S.A.H. (Eds.). Biodiversidade e ecossistemas bentônicos marinhos do Litoral Norte de São Paulo, Sudeste do Brasil. Campinas, SP, UNICAMP. 573p.
Barattini, L.P. & Ureta, E.H. 1961. La fauna de las costas uruguayas del este (invertebrados). Montevideo, Museo Damaso Larrañaga. 196p., 52 pls. (Publicación de Divulgación Cientifica, Museo “Damaso Larrañaga”, Montevideo).
Barroso, C.X.; Rabay, S.G.; Meirelles, C.A.O. & Matthews-Cascon, H. 2013. Mollusks from two estuarine areas in Ceará State, northeastern Brazil, with new state records for four species. Check List, 9(3): 504-509. https://doi.org/10.15560/9.3.504.
Bernard, F.R.; Cai, Y.Y. & Morton, B. 1993. Catalogue of the living Marine Bivalve Molluscs of China. Hong Kong, Hong Kong University Press.121p. https://doi.org/10.1515/9789882200555.
Bertin, V. 1876. Révision des Tellinidés du Museum d'Histoide Naturelle. Nouvelles Archives du Muséum d'Histoire Naturelle (ser. 2), 1: 201-361, pl. 8-9.
Bieler, R. & Mikkelsen, P.M. 2025. A taxonomic update for “Seashells of Southern Florida – Bivalves” (Mikkelsen and Bieler, 2007) with nomenclatural notes on other bivalve species. Nautilus, 139: 1-27. https://doi.org/10.1515/9780691239453.
Bonanni, P. 1684. Recreatio mentis, et oculi in obseruatione animalium testaceorum. Rome, Varesij. 207p.; 404 ilus.
Born, I. 1780. Testacea Musei Caesarei Vindobonensis. Vindobonae, Joannis Pauli Kraus. 442p., 18 pls. https://doi.org/10.5962/bhl.title.152209.
Bory de Saint-Vincent, J.B.M. 1827. Tableau encyclopédique et méthodique des trois règnes de la nature vers, coquilles, mollusques et polypiers tome premier. Paris, Agasse. 180p.; pls. 1-95.
Borzone, C.A.; Vargas, K.M.; Pezzuto, P.R. & Tavares, Y.A.G. 2001. Aspectos da reprodução e dinâmica populacional de Chione pubera (Bory Saint-Vicent) (Bivalvia, Veneridae) no sul do Brasil. Revista Brasileira de Zoologia, 18(2): 333-349. https://doi.org/10.1590/S0101-81752001000200003.
Bosc, L.A.G. 1824. Histoire Naturelle des Coquilles. Paris, Chez Verdière. 287p.
Boss, K.J. 1966. The subfamily Tellininae in the Western Atlantic the genus Tellina. Johnsonia, 4(45-47): 217-364.
Boss, K.J. & Kenk, V.C. 1964. Anatomy and Relationships of Temnoconcha brasiliana Dall. Occasional Papers on Mollusks, 2(30): 325-343.
Boss, K.J.; Rosewater, J. & Ruhoff, F.A. 1968. The zoological taxa of William Healey Dall. Bulletin of the United States National Museum, 287: 1-427. https://doi.org/10.5479/si.03629236.287.
Bouchet, P. 1997. Inventorying the molluscan diversity of the world: what is our rate of progress? Nautilus, 40: 1-11.
Bouchet, P.; Bary, S.; Héros, V. & Marani, G. 2016. How many species of molluscs are there in the world’s oceans, and who is going to describe them? In: Héros, V.; Strong, E. & Bouchet, P. (Eds.). Tropical Deep-Sea Benthos, 29. Muséum national d’Histoire naturelle, Paris: 9-24 (Mémoires du Muséum National d’Histoire Naturelle, 208) ISBN: 978-2-85653-774-9.
Brann, D.C. & Kent, L.S. 1960. Catalogue of the type and figured specimens in the Paleontological Research Institution. Bulletins of American Paleontology, 40(184): 1-995.
Bruguière, J.G. 1791. Tableau Encyclopédique et Méthodique des trois Règnes de la Nature: vers, coquilles, mollusques et polypiers divers. Paris, Panckoucke. 1: 1-180, 92 pls. https://www.biodiversitylibrary.org/item/115590#page/7/mode/1up.
Bruguière, J.G. 1797. Tableau Encyclopédique et Méthodique des trois Règnes de la Nature: dix-neuvième partie. Vers testacées, acoquilles bivalves. Henri Agasse. Paris, 18: pls. 93-286. https://www.biodiversitylibrary.org/item/156156#page/9/mode/1up.
Buckup, L. & Buckup, E.H. 1957. Catálogo dos moluscos do Museu Rio-Grandense de Ciências Naturais. Iheringia Série Zoologia, 1: 1-40.
Cardoso, P.S. & Rios, E.C. 1967. Lista preliminar de los moluscos marinos de Alagoas. Comnicaciones de la Sociedad Malacologica del Uruguay, 2(13): 117-135. https://doi.org/10.5962/p.421026.
Carpenter, P.P. 1857. Report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science, 26: 159-368, pls. 6-9. https://doi.org/10.5962/bhl.title.60613.
Castellanos, Z.J.A. 1970. Catálogo de los moluscos marinos bonaerenses. Anales de la Comisión de Investigaciones Científicas de la Provincia de Buenos Aires, 8: 9-365.
Catullo, T.A. 1828. Zoologie du calcaire jurassique de l’Apennis. Bulletin des Sciences Naturelles et de Géologie, 13: 250-252.
Cauquoin, M. 1967. Lamellibranchiate molluscs – Tellinidae, Scrobiculariidae and Donacidae. Annales de l’Institut Oceanographique, 45(2): 227-223. https://doi.org/10.3406/adh.1967.960.
Chemnitz, J.H. 1782. Neues systematisches Conchylien-Cabinet. Gabriel Nicolaus Raspe. Nürnberg, 6: 375 pp., pls. 1-36. https://doi.org/10.5962/bhl.title.120076.
Chenu, J.C. 1844. Illustrations conchyliologiques, ou description et figures de toutes les coquilles connues, vivantes et fossiles. Spondylus. Spondyle, Linné. Paris. 9p. pl. 31-32. https://www.biodiversitylibrary.org/item/218318#page/285/mode/1up
Chuang, S.H. 1973. Animal life and nature in Singapore. Singapore University Press. 302p.
Coan, E.V. 1997. Recent species of the genus Petricola in the eastern Pacific (Bivalvia: Veneroidea). Veliger, 40(4): 298-340.
Coan, E.V. 1999. The eastern Pacific species of Sphenia (Bivalvia: Myidae). Nautilus, 113(4): 103-120. https://doi.org/10.5962/bhl.part.2019.
Coan, E.V. & Valentich-Scott, P. 2008. Three nomenclatural notes on Panamic bivalves. Festivus, 40(4): 49-54.
Coltro, J. 1997. 15 years collecting around São Sebastião, São Paulo, Brazil. American Conchologist, 25: 7-11.
Coltro, J. 1998. The Brazilian connection: let’s go shelling in Brazil – Part 1. American Conchologist, 26: 7-9.
Cosel, R.v. 1986. Moluscos de la region de la Cienaga Grande de Santa Marta (Costa del Caribe de Colombia). Anales del Instituto de Investigaciones Marinas de Punta de Betin, 15-16: 79-370.
Cosel, R.v. & Gofas, S. 2019. Marine bivalves of tropical West Africa: from Rio de Oro to southern Angola. Faune et Flore Tropicales, 48: 1-1104. https://doi.org/10.5852/fft48.
Cosel, R.v.; Gofas, S. & Poutiers, J.M. 2014. Nomenclatural notes on some European and West African marine bivalve species. Iberus, 32: 65-85.
Cox, L.R. 1929. On the varietal names for Spondylus in K. Schreibers’ Versuch einer Vollstãndigen Conchylienkenntniss (1793). Proceedings of the Malacological Society of London, 18: 251-254. https://doi.org/10.1093/oxfordjournals.mollus.a063975.
Crouch, C. 2026. Taxonomy & nomenclature. Website. https://sites.broward.edu/science-wellness-central/physical-sciences/paleo/taxonomy. Access: 16/05/2026.
d’Orbigny, C.D. (Ed.). 1849. Dictionnaire universel d’histoire naturelle. Atlas 3, 1-27, 35 pls. https://www.biodiversitylibrary.org/item/272842#page/9/mode/1up.
Daccarett, E.Y. & Bossio, V.S. 2011. Colombian Seashells from the Caribbean Sea. L’Informatore Piceno. 384p.
Dall, W.H. 1902. Synopsis of the family Veneridae and of the North American Recent species. Proceedings of the United States National Museum, 26(1312): 335-412, pl. 12-16. https://doi.org/10.5479/si.00963801.26-1312.335.
Dall, W.H. 1903. Contributions to the Tertiary fauna of Florida. Transactions of the Wagner Free Institute of Science of Philadelphia, 3(4): 1219-1654.
Dall, W.H. 1921. Two new South American shells. Nautilus, 34(4): 132-133. https://doi.org/10.1002/ange.19210342706.
Dall, W.H. & Simpson, C.T. 1901. The Mollusca of Porto Rico. Bulletin of the United States Fish Commission, 20: 351-524, pls. 53-58.
Dance, S.P. 1974. The collector’s encyclopedia of shells. New York, McGraw-Hill Book Company. 288p.
Davenport, R. 1995. What I did this summer. Texas Conchologist, 32(1): 9-15.
Davies, F.R.E. 1991. Bellow Capricorn. American Conchologist, 17(2): 22.
Deshayes, G.P. 1853. Catalogue of the Conchifera or bivalve shells in the collection of the British Museum. Part I. Veneridae, Cyprinidae and Glauconomidae. London, British Museum. 216p.
Díaz, J.M. & Puyana, M. 1994. Moluscos del Caribe Colombiano. Bogota, Invemar. 292p., 74 pls. (Colciencias. Fundación Natura)
Dillwyn, L.W. 1817. A descriptive catalogue of recent shells. Cornhill, John and Arthur Arch. 1-2: xii, 1121p.
Domaneschi, O. 1984. Família Myidae Lamarck, 1809. Informativo da Sociedade Brasileira de Malacologia, 40: 9-12.
Domaneschi, O. & Narchi, W. 1998. Adaptive convergences in two nestling bivalves (Myoidea: Myidae, Hiatellidae) of the Brazilian littoral. Iheringia Série Zoologia, 85: 89-96.
Dornellas, A.P.S. & Simone, L.R.L. 2011. Annotated list of types specimens of molluscks deposited in Museu de Zoologia da Universidade de São Paulo, Brazil. Arquivos de Zoologia, 42(1): 1-81. https://doi.org/10.11606/issn.2176-7793.v42i1p1-81.
Espinosa, E. & Ortea, J. 2001. Moluscos del Mar Caribe de Costa Rica: desde Cahuita hasta Candoca. Avicennia, suppl. 4: 1-77.
Espinosa, E.; Ortea, J. & Caballer, M.M.L. 2006. Marine Mollusca from Guanahacabibes peninsula, Pinar del Rio, Cuba, with description of new taxa. Avicennia, 18: 1-84.
Espinosa, E.; Ortea, J. & Valdés, A. 1994. Catálogo de los moluscos bivalvos recientes del Archipiélago cubano. Avicennia, 2: 109-129.
Espinosa, J.; Ortea, J.; Sánchez, R. & Gutiérrez, J. 2012. Moluscos marinos Reserva de la Biosfera de la Península de Guanahacabibes. Instituto de Oceanología. La Habana, 325 pp.
Favanne, M. & Favanne, J. 1780. La conchyliologie, ou, Histoire naturelle des coquilles de mer, d’eau douce, terrestres et fossiles: avec un traité de la zoomorphose, ou, représentation des animaux qui les habitent, ouvrage dans lequel on trouve une nouvelle méthode de les diviser. Paris. 80 plates. https://www.biodiversitylibrary.org/item/81331#page/1/mode/1up.
Finet, Y. & Lamprell, K.L. 2008. The Spondylidae in the historical collections of the Muséum d’Histoire Naturelle, Geneva, with revision of the species described by Lamarck and by Chenu, and notes on nomenclature. Archiv für Molluskenkunde, 137: 1-73. https://doi.org/10.1127/arch.moll/0003-9284/137/001-073.
Fischer, G. 1808. Tableaux synoptiques de Zoognosie. Moscou, Universitè Impériale. 186p.
Fischer-Piette, E. 1975. Révision des Veneridae s.s. (mollusques lamellibranches). Mémoires du Muséun National d’Histoire Naturelle, n.s. A-Zoologie, 93: 1-64, pls. 1-8.
Fischer-Piette, E. & Lamy, E. 1942. Notes sur les Veneridæ (Moll. Lamellibr.) de Quoy et Gaimard. Mémoires du Muséum National d’Histoire Naturelle, ser. 2, 14: 130-137.
Fischer-Piette, E. & Testud, A.M. 1967. Mollusques Lamellibranches: Veneridae. In: Forest, J. (Ed.). Résultats Scientifiques des Campagnes de la Calypso 8. Annales de l’Institut Océanographique, 45: 205-220.
Fischer-Piette, E. & Testud, A.M. 1970. Veneridae du Surinam communiquées par M.C.O., van Regteren Altena. Bulletin du Muséum Nationa d’Histoire Naturelle, ser. 2, 42(3): 520-524.
Fischer-Piette, E. & Vukadinovic, D. 1977. Suite des revisions des Veneridae (Moll. Lamellibr.) Chioninae, Samaranginae et complément aux vénus. Mémoires du Muséum National d’Histoire Naturelle, n.s., Zoologie, 106: 1-186, 22 pls.
Fischer-Piette, E.; Kempf, M. & Testud, A.M. 1970. Nouvelles données sur Ies Veneridae (Mollusques Lamellibranches) du Brésil. Bulletin du Museum d’Histoire Naturelle, 41: 1543-1553.
Folin, L. 1867. Les méléagrinicoles. Espèces nouvelles. Recueil des Publications Societé Havraise d’Étude Diverse, 33: 41-112, 6 pls. https://doi.org/10.5962/bhl.title.10790.
Folin, L. & Périer, L. 1867. Les fonds de la mer. Etude internationnale sur les particularités nouvelles des régions sous-marines. Paris, Boulevard Lib. Boulevard Saint Germain. v. 1, 48p.
Forti, I.R.S. 1969. Cenozoic mollusks from the drill-holes Cassino and Palmares do Sul of the coastal plain of Rio Grande do Sul. Iheringia Série Geologia, 2: 55-155.
Fulton, H.C. 1937. Notes on the Recent Spondylus of Florida. Nautilus, 51(2): 38-39.
Galli, C. 2024. WMSDB – Worldwide Mollusc Species DataBase. Genova, Italy. Website. https://www.conchology.be/?t=2203&family=&Genus=anomalocardia&species=&variety=&AUTHOR=. Access: 29/07/2024.
García, E.F. & Lee, H.G. 2001. Report on molluscan species found in the offshore waters of Lousiana, including many extensions of known range and un-named species. American Conchologist, 30: 10-13.
Geraldes, 1982. Contribuição ao conhecimento de Macoma (Austromacoma) constricta (Bruguière, 1792) (Bivalvia, Heterodonta, Tellinidae). M.Sc. Dissertation. Rio de Janeiro, Univerdidade Federal do Rio de Janeiro. 105p. https://pantheon.ufrj.br/bitstream/11422/2740/1/278283.pdf.
Gmelin, J.F. 1791. Systema naturae per regna tria naturae. Editio decima tertia. Lipsiae, v. 1, p. 021-3910.
Gray, J.E. 1853. Calatogue of conchifera or bivalve shells in the collection of the British Museum. London. 216p.
Gualtieri, N. 1742. Index testarum conchyliorum quae adservantur in Museo N. Gualtieri … et methodice distributae exhbentur tabulis ex. Florentinae. 5 pt, 110 pls. https://doi.org/10.5962/bhl.title.132271.
Haburay, K. 1976. New locality records for Spirobranchus giganteus var. giganteus in the northeastern Gulf of Mexico. Florida Scientist, 39(2): 127-128.
Hanley, S.C.T. 1855. The shells of Linnaeus, determined from his manuscripts and collection. London, Ipsa Linnaei Conchylia. 556p. https://doi.org/10.5962/bhl.title.11868.
Hartmann, T. 2006. Bivalve seashells of Florida. Korea, Anadara Press. 183p.
Haviland, E. 1990. Records of sea molluscs of Brazil much to do about Spondylidae. Siratus, 7: 3-6, 11.
Hebard, E.B. 1967. Pleistocene mollusks from New Providence Island, Bahamas. Nautilus, 81: 41-44.
Heise, J.R. 2024. Temnoconcha brasiliana Dall, 1921. Conquiliologistas do Brasil. Espécies do Brasil. Website: http://www.conchasbrasil.org.br/conquiliologia/descricao.asp?id=347. Access: 27/04/2024.
Hertlein, L.G. & Strong, A.M. 1948. Eastern Pacific expeditions of the New York Zoological Society. XXXIX. Mollusks from the west coast of Mexico and Central America. Part VI. Zoologica, 33(4): 163-198, 2 pls. https://doi.org/10.5962/p.184650.
Hewitt, S.J. 2010. A three-minute survey of the marine mollusks of the island of Sint Eustatius, Leeward Islands, West Indies. Festivus, 42: 35-38.
Hewitt, S.J. 2013. A second sort survey of the marine mollusks of the Island Montserrat, Leeward Islands, West Indies. Festivus, 45(11): 97-103.
Hidalgo, J.G. 1903. Obras malacológicas. Parte 1, Islas Filipinas. Madrid, Imprenta de L. Aguado. 400p.
Huber, M. 2010. Compendium of bivalves. A full-color guide to 3,300 of the world’s marine bivalves. A status on Bivalvia after 250 years of research. Hackenheim, ConchBooks. 901p.
Huber, M. 2015. Compendium of bivalves 2. Harxheim, ConchBooks. 907p.
Huber, M.; Langleit, A. & Kreipl, K. 2015. Tellinidae; In: Huber, M. Compendium of bivalves 2. Harxheim, ConchBooks. p. 564-745.
Hylleberg, J. & Kilburn, R.N. 2003. Marine Molluscs of Vietnam. Phuket Marine Biological Center Special Publication, 28: 5-300.
Ihering, H.v. 1907. Les Mollusques fossiles du Tertiare et du Crétacé Supérieur de l’Argentine. Anales del Museo Nacional de Buenos Aires, (3)7: 1-611, 18 pls.
International Commission on Zoological Nomenclature (ICZN) 1999. International Code of Zoological Nomenclature, 4. ed. London,International Trust for Zoological Nomenclature. 306p. https://www.iczn.org/the-code/the-code-online. Access: 03/04/2025.
Jay, J.C. 1850. A catalogue of the shells, arranged according to the Lamarckian system: with their authorities, synonymes, and references to works where figured or described, contained in the collection of John C. Jay. Fourth edition. New York, Craighead. 460p. https://doi.org/10.5962/bhl.title.13211.
Jay, J.C. 1852. Supplement. In: A Catalogue of the shells in the collection, arranged according to the Lamarkian system with a supplement containing additional species, corrections, and alterations of John C. Jay. 4. ed. New York. p. 465-479. https://www.biodiversitylibrary.org/item/70818#page/477/mode/1up.
Jones, C.C. 1979. Anatomy of Chione cancellata and some other chionines (Bivalvia: Veneridae). Malacologia, 19(1): 157-199.
Keen, A.M. 1971. Sea shells of tropical West America, second edition. Stanford, Stanford University Press. 1064p., 22 pls.
Kinkelin, F. 1903. Die Originale der paläontologischen Sammlung im Senckenbergischen Museum und die auf dieselben bezügliche Literatur. Bericht der Senckenbergischen Naturforschenden Gesellschaft, 1903, 3-154.
Knight, R. & Morris, N.J. 2019. Well-developed muscle attachments in British Albian inoceramids (Inoceramidae, Bivalvia): implications for inoceramid palaeobiology, evolution and taxonomy. Papers in Palaeontology, 5: 1-21. https://doi.org/10.1002/spp2.1246.
Koch, A. 1876. Geologisch Beschaffenheit der am rechten Ufer gelegenen Hälfte der Donautrachytgruppe*) (Sct. Andrä-Visegrader Gibirgsstock) nahe Budapest. Zeitschrift der Deutschen Geologischen Gesellschaft, 28: 293-349.
Lamarck, J.B.P.M. (by Deshayes, G.P.) 1835. Histoire naturelle des animaux sans vertèbres. Paris, J.B. Baillière. v. 2, pt. 6, 600p. https://doi.org/10.5962/bhl.title.2836.
Lamarck, J.B.P.M. (by Deshayes, G.P.), 1838. Histoire naturelle des animaux sans vertèbres. 10. ed. Paris, J.B. Baillière. v. 9, 728 pp. https://doi.org/10.5962/bhl.title.4846.
Lamarck, J.B.P.M. 1818. Histoire naturelle des animaux sans vertèbres. Paris. v. 5, 612p.
Lamprell, K.L. 1989. Spondylus – Notes on the American species. American Conchologist, 17(1): 6-9.
Lamprell, K.L. 2006. Spiny oysters: a revision of the living Spondylus species of the world. Brisbane. 119p.
Lamy, D.; Massemin, D, Pointier, J.P. & Gargominy, O. 2009. Coquillages et escargots de Guyane. Paris, Editions Biotope, Mèze and Muséum National d’Histoire Naturelle. 456p.
Lamy, E. 1938. Révision des Spondylus vivants du Muséum National d’Histoire Naturelle de Paris. Journal de Conchyliologie, 82: 265-306.
Lamy, E. & Fischer-Piette, E. 1938. Notes sur les espèces Lamarckiennes de Venus s. str. et de Chione (Moll. Lamellibr.). Bulletin du Museum National d’Histoire Naturelle, Series 2, 10(4): 401-405.
Lima, V.B. 2017. Riqueza de moluscos associados aos recifes do Morro de São Paulo, Ilha Tinharé, Bahia (nordeste do Brasil). Campina Grande, Universidade Federal de Campina Grande. 27p.
Linné, C. 1767. Systema Naturae. 12. ed. Holmiae, Laurentii Salvii. v. 1, pt. 2, p. 533-1327.
Lister, M. 1685-1692. Historiae methodicae conchyliorum … liber primus – liber iv. – Appendix ad … librum iv – [Tabularum anatomicarum explicatio]. Londini, 6 pt, 2 vols.
Long, S. 1985. Suggested list of common names for mollusks occurring from America Borth of Mexico. Shells and Sea Life, 9: 45-85.
Lopes, R.P. & Simone, L.R.L. 2012. New fossil records of Pleistocene marine mollusks in southern Brazil. Revista Brasileira de Paleontologia, 15(1): 49-56. https://doi.org/10.4072/rbp.2012.1.04.
Lynge, H. 1909. The Danish expedition to Siam 1899-1900. IV. Marine Lamellibranchiata. Det Kongelige Danske Videnskabernes Selskabs Medlemmer, Naturvidenskabelig og Mathematisk Afdeling, (7)5(3): 97-299.
Macedo, I. 2013. A comunidade macrobentônica da endofauna na região sob influência da desembocadura do Rio Paraíba do Sul. Rio de Janeiro, Universidade Estadual do Norte Fluminense. 75p.
Marché, T. & Grace, T. 1992. Shelling the ghost fleet of the Atlantic. American Conchologist, 20: 8-11.
Marques, R.C. & Simone, L.R.L. 2014. Eurytellina angrensis, a new species from southeastern Brazilian coast, with anatomical details (Bivalvia: Tellinoidea). Archiv für Molluskenkunde, 143(1): 39-50. https://doi.org/10.1127/arch.moll/1869-0963/143/039-050.
Marques, R.C.; Silva, A.M. & Simone, L.R.L. 2022. Cladistic analysis of the transisthmian genus Eurytellina (Bivalvia: Tellinoidea) based on morphological and morphometric data. Organisms Diversity & Evolution, 22: 857-891. https://doi.org/10.1007/s13127-022-00561-z.
Marshall, B.A. & Spencer, H.G. 2013. Comments on some taxonomic changes affecting marine Bivalvia of the New Zealand region recently introduced in Huber’s Compendium of bivalves, with some additional taxonomic changes. Molluscan Research, 33(1): 44-49. https://doi.org/10.1080/13235818.2012.754147.
Martínez, S. & Del Río, C.J. 2005. Las Ingresiones marinas del Neógeno en el sur de Entre Ríos (Argentina) y Litoral Oeste de Uruguay y su contenido malacológico. In: Aceñolaza, F.G. (Org.). Temas de la Biodiversidad del Litoral fluvial argentino II. INSUGEO Miscelánea, 14: 13-26.
Maury, C.J. 1925. A further contribution of the paleontology of Trinidad (Miocene horizons). Bulletin of American Paleontology, 10(42): 153-404, 12-54 pls.
McLean, R.A. 1951. The Pelecypoda or bivalve mollusks of Porto Rico and the Virgin Islands. Scientific Survey of Porto Rico and the Virgin Islands, 17(1): 1-183.
Mikkelsen, P.M. & Bieler, R. 2004. Critical catalog and annotated bibliography of marine bivalve records for the Florida Keys. Malacologia, 46: 545-623.
MolluscaBase (Eds.). 2026. MolluscaBase. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=533480. Access: 16/05/2026.
Mont, M.A.; Whyman, J. & Khlopas, A. 2020. The Atlantic thorny oyster. Molluscan Science – A World of Fascination and Beauty, 1: 1-43.
Morgan, K.; Lloyd, C. & Chester, J. 1997. Red cones at night … Shellers’ delight. American Conchologist, 27: 4-8, 15.
Morretes, F.L. 1949. Ensaio de catálogo dos moluscos do Brasil. Arquivos do Museu Paranaense, 7: 1-216.
Nabhitabhata, J. 2009. Checklist of Mollusca fauna in Thailand. Bangkok, Office of Natural Resources and Environmental Policy and Planning. 576p.
Narchi, W. 1975. Functional morphology of a new Petricola (Mollusca Bivalvia) from the littoral of Sao Paulo, Brazil. Proceedings of the Malacological Society of London, 41: 451-465.
Narchi, W. & Domaneschi, O. 1993. The functional anatomy of Sphenia antillensis Dall & Simpson, 1901 (Bivalvia: Myidae). Journal of Molluscan Studies, 59(2): 195-210. https://doi.org/10.1093/mollus/59.2.195.
Odé, H. & Speers, A.B. 1967. Notes concerning Texas beach shells [Family Veneridae]. Texas Conchologist, 4-5: 50-51.
Oliveira, G.S.P. 1994. (“1992”) Moluscos en sedimentos biogênicos da Ponta de Itapuã, Salvador, Bahia (1). Comunicaciones de la Sociedad Malacológica del Uruguay, 7(62-63): 277-303. https://doi.org/10.5962/p.421204.
Olsson, A.A. 1961. Mollusks of the tropical eastern Pacific, particularly from the southern half of the Panamic-Pacific faunal province (Panama to Peru). Panamic-Pacific Pelecypoda. Paleontological Research Institution. Ithaca, 574p., pls. 1-86. https://doi.org/10.5962/bhl.title.6853.
Palacio, E.F.; Gracia, A. & Duque, G. 2010. Benthic Mollusks of La Guajira (10 and 50 m depths), Colombian Caribbean. Boletín de Investigaciones Marinas y Costeras-INVEMAR, 39(2): 397-416.
Palmer, K.E.H.v.W. 1927. The Veneridae of eastern North America, Cenozoic and Recent. Palaeontographica Americana, 1(5): 209-522, pls. 32-76.
Passos, F.D. & Magalhães, F.T. 2011. A comparative study of the Bivalvia (Mollusca) from the continental shelves of Antarctica and Brazil. Biota Neotropica, 11(1): 143-155. https://doi.org/10.1590/S1676-06032011000100014.
Passos, F.D.; Bastião, A.R. & Cardoso, L.L. 2024. Checklist of marine Bivalvia (Mollusca) from Brazil, with descriptive analyses of their bathymetric and geographical distribution. Zootaxa, 5488(1): 1-94. https://doi.org/10.11646/zootaxa.5488.1.1.
Pastorino, G. & Bagur, M. 2011. The genus Sphenia Turton, 1822 (Bivalvia: Myidae) from shallow waters of Argentina. Malacologia, 54(1-2): 431-435. https://doi.org/10.4002/040.054.0112.
Perry, G. 1811. Conchology, or the natural history of shells. London, William Miller. 255p., 61 pls.
Perry, L.M. 1940. Marinae shells of the southwest coast of Florida. Bulletins of American Paleontology, 26: 1-260.
Perry, L.M. & Schwengel, J.S. 1955. Marine shells of the western coast of Florida. Ithaca, Paleontological Research Institution. 318p. https://doi.org/10.5962/bhl.title.6586.
Petuch, E.J. & Myers, R.F. 2014. Molluscan communities of the Florida Keys and adjacent areas. Their ecology and biodiversity. Boca Raton, CRC Press. 300p. https://doi.org/10.1201/b17583.
Piffer, P.R.; Arruda, E.P. & Passos, F.D. 2011. The biology and functional morphology of Macoma biota (Bivalvia: Tellinidae: Macominae). Zoologia, 28(3): 321-333. https://doi.org/10.1590/S1984-46702011000300006.
Pilsbry, H.A. 1897. New species of mollusks from Uruguay. Proceedings of the Academy of Natural Sciences of Philadelphia, 49: 290-298, pls. 6-7.
Pilsbry, H.A. 1899. Littoral mollusks from cape Fairweather, Patagonia. American Journal of Science (ser. 4), 7: 126-128. https://doi.org/10.2475/ajs.s4-7.38.126.
Prado, A.C.G. & Abreu, O.S. 2000. Conchas brasileiras – Lista de Recordes II / Brazilian Shells – Record List II. Strombus, 7: 1-12.
Prado, A.C.G. & Abreu, O.S. 2001. Conchas brasileiras – Lista de Recordes III / Brazilian Shells – Record List III. Strombus, 8: 1-14.
Pulley, T.E. 1952. An illustrated checklist of the marine mollusks of Texas. Texas Journal of Science, 4: 167-199.
Reeve, L. 1856. Monograph of the genus Spondylus. Conchologia Iconica, 9: pls. 1-18.
Reeve, L. 1863. Monograph of the genus Venus. Conchologia Iconica, 14: pls. 1-26.
Reeve, L. 1870. Monograph of the genus Tellina. Conchologia Iconica, 17: pls. 1-58.
Richards, H.G. 1953. Pleistocene mollusks from Andros Island, Bahamas. Nautilus, 67(4): 120-121.
Rios, E.C. 1970. Coastal Brazilian seashells. Rio Grande, Fundação Cidade do Rio Grande. 255p., 4 maps, 60 pls.
Rios, E.C. 1975. Brazilian marine mollusks iconography. Rio Grande, Fundação Cidade do Rio Grande. 331p., 91 pls.
Rios, E.C. 1985. Seashells of Brazil. Rio Grande, Museu Oceanográfico. 329p., 102 pls.
Rios, E.C. 1994. Seashells of Brazil, second edition. Rio Grande, Fundação Universidade do Rio Grande. 368p., 113 pls.
Rios, E.C. 2009. Compendium of Brazilian sea shells. Rio Grande, Universidade Federal do Rio Grande. 668p.
Rocha, V.P. & Matthews-C., H. 2015. Inferência de padrões de distribuição da família Veneridae (Mollusca, Bivalvia) no Brasil através de base secundária de dados. Neotropical Biology and Conservation, 10(3): 123-131. https://doi.org/10.4013/nbc.2015.103.02.
Rocha, V.P.; Silveira, I.O. & Matthews-C., H. 2015. Brazilian Spondylidae: a brief discussion about variation of shell ornamentation in the northeastern species. Arquivos de Ciências do Mar, 48: 79-84.
Rosenberg, G. 2011. Malacolog Version 4.1.1. A Database of Western Atlantic Marine Mollusca. Philadelphia [website http://www.malacolog.org].
Rumphius, G.E. 1741. D’Amboinsche Rariteitkamer. Jan Roman de Jonge. Amsterdam, 340 pp, 60 pls. https://www.biodiversitylibrary.org/item/127485#page/11/mode/1up.
Sandberger, C.L.F. 1863. Mainzer Tertiärbeckens. Wiesbaden, C.W. Kreidel’s Verlag. 458p.
Scarabino, F. & Zaffaroni, J.C. 2004. Estatus faunístico de veinte especies de moluscos citadas para águas uruguayas. Comunicaciones Zoologicas del Museo Nacional de Historia Natural y Antropología, 8(202): 1-16.
Scarabino, F.; Zelaya, D.G.; Orensanz, J.L.; Ortega, L.; Defeo, O.; Schwindt, E.; Carranza, A.; Zaffaroni, J.C.; Martínez, G.; Scarabino, V. & García-Rodríguez, F. 2016. Cold, warm, temperate and brackish: Bivalve biodiversity in a complex oceanographic scenario (Uruguay, southwestern Atlantic). American Malacological Bulletin, 33(2): 284-301, suppl. mat, 1-35. https://doi.org/10.4003/006.033.0219.
Schreibers, K.v. 1793. Versuch einer vollständigen Conchylienkenntniss nach Linnes System. Wien, Joseph Edlen v. Kurzbeck. 447p. https://doi.org/10.5962/bhl.title.35864.
Sistema de Informação sobre a Biodiversidade Brasileira (SIBBr). 2024. Sistema de informação sobre a biodiversidade brasileira. Website: https://ala-bie.sibbr.gov.br/ala-bie/species/225593. Access: 27/07/2024.
Silveira, I.O. 2017. Morfologia de espécies do gênero Spondylus (Mollusca: Bivalvia, Spondylidae) pepositados na coleção malacológica Prof. Henry Ramos Mattews. Fortaleza,Universidade Federal do Ceará. 28p.
Simone, L.R.L. 2022. The molluscan jugal muscles – m1. Malacopedia, 5(5): 43-48. https://www.moluscos.org/trabalhos/Malacopedia/05-05Simone%202022%20Malacopedia-jugal%20muscle%20m1.pdf.
Simone, L.R.L.; Mikkelsen, P.M. & Bieler, R. 2015. Comparative anatomy of selected marine bivalves from the Florida Keys, with notes on Brazilian congeners (Mollusca: Bivalvia). Malacologia, 58(1-2): 1-127. https://doi.org/10.4002/040.058.0201.
Smith, M. 1937. East coast marine shells: descriptions of shore mollusks together with many living below tide mark, from Maine to Texas inclusive, especially Florida, with more than one thousand drawings and photographs. Lantana. 314p. https://doi.org/10.5962/bhl.title.6796.
Sowerby II, G.B. 1847a. Descriptions of several new species of Spondylus. Proceedings of the Zoological Society of London, 15(175): 86-88.
Sowerby II, G.B. 1847b. Monograph of the genus Spondylus. In: Thesaurus Conchyliorum monographs of genera of shells v. 1, p. 417-438, pls. 85-89.
Sowerby II, G.B. 1853. Monograph of the genus Venus: supplement. In: Thesaurus Conchyliorum monographs of genera of shells. v. 2, p. 703-762, pls. 152-163.
Sowerby II, G.B. 1868. Monograph of the genus Tellina. Conchologia Iconica London, Reeve. pt. 17, 58 pls. https://www.biodiversitylibrary.org/item/41303#page/161/mode/1up
Tallarico, L.F.; Passos, F.D.; Machado, F.M.; Campos, A.; Pimental, S.M.R. & Introíni, G.O. 2014. Bivalves of the São Sebastião Channel, north coast of the São Paulo State, Brazil. Check List, 10(1): 97-105. https://doi.org/10.15560/10.1.97.
Tenorio, D.O. 1984. O gênero Tellina Linnaeus, 1758 (Molusca[sic], Bivalvia) na plataforma continental brasileira. Trabalhos Oceanográficos da Universidade Federal de Pernambuco, 18: 7-138. https://doi.org/10.5914/tropocean.v18i1.2608.
Turgeon, D.D.; Lyons, W.G.; Mikkelsen, P.; Rosenberg, G. & Moretzsohn, F. 2009. Bivalvia (Mollusca) of the Gulf of Mexico. In: Felder, D.L. & Camp, D.K. (Eds.). Gulf of Mexico-Origins, Waters, and Biota. Biodiversity. Texas A&M Press, Colleg. p. 711-744
VanderVen, K. 2011. Romancing the stones: a shelling expedition to the Dominican Republic. American Conchologist, 39: 15-20.
Viana, M.G. & Rocha-B., C.A. 2007. The sensorial Structures of Spondylus americanus Hermann, 1781 (Mollusca: Bivalvia, Spondylidae). Brazilian Archives of Biology and Technology, 50(5): 815-819. https://doi.org/10.1590/S1516-89132007000500009.
Vilvens, C. & Meuleman, E. 2008. Queleues belles coquilles d’Amerique du Sud. Novapex, 9: 29-33.
Warmke, G.L. & Abbott, R.T. 1961. Caribbean seashells. Dover Publication, Inc. New York. 348p.
Wells, F.E.; Sanpanich, K.; Tan, S.K. & Duangdeem, T. 2021. The marine and estuarine molluscs of Thailand. Singapore, Lee Kong Chian Natural History Museum. 195p.
Downloads
Published
Issue
Section
License
Copyright (c) 2026 Luiz Ricardo L. Simone, Fabrizio M. Machado
This work is licensed under a Creative Commons Attribution 4.0 International License.
Responsibility: The scientific content and the opinions expressed in the manuscript are the sole responsibility of the author(s).
Copyrights: The Authors. The journal is licensed under a Creative Commons attribution-type CC-BY.
How to Cite
Funding data
-
Conselho Nacional de Desenvolvimento Científico e Tecnológico
Grant numbers 301717/2022-8 -
Fundação de Amparo à Pesquisa do Estado de São Paulo
Grant numbers 2014/10483-1;2010/11400-1
